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Backward-stochastic-differential-equation approach to modeling of gene expression

Evelina Shamarova1,*, Roman Chertovskih2, Alexandre F. Ramos3, and Paulo Aguiar4

  • 1Departamento de Matemática, Universidade Federal da Paraíba, 58051-900 João Pessoa, Brazil
  • 2Samara National Research University, Moskovskoe shosse 34, 443086 Samara, Russian Federation
  • 3Escola de Artes, Ciências e Humanidades, Universidade de São Paulo, Av. Arlindo Béttio 1000, 03828-00 São Paulo, SP, Brazil
  • 4INEB, Instituto de Engenharia Biomédica i3S, Instituto de Investigação e Inovação em Saúde, Rua Alfredo Allen 208, 4200-135 Porto, Portugal

  • *evelina@mat.ufpb.br

Phys. Rev. E 95, 032418 – Published 29 March, 2017

DOI: https://doi.org/10.1103/PhysRevE.95.032418

Abstract

In this article, we introduce a backward method to model stochastic gene expression and protein-level dynamics. The protein amount is regarded as a diffusion process and is described by a backward stochastic differential equation (BSDE). Unlike many other SDE techniques proposed in the literature, the BSDE method is backward in time; that is, instead of initial conditions it requires the specification of end-point (“final”) conditions, in addition to the model parametrization. To validate our approach we employ Gillespie's stochastic simulation algorithm (SSA) to generate (forward) benchmark data, according to predefined gene network models. Numerical simulations show that the BSDE method is able to correctly infer the protein-level distributions that preceded a known final condition, obtained originally from the forward SSA. This makes the BSDE method a powerful systems biology tool for time-reversed simulations, allowing, for example, the assessment of the biological conditions (e.g., protein concentrations) that preceded an experimentally measured event of interest (e.g., mitosis, apoptosis, etc.).

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